Gramma the Galápagos tortoise died on Nov. 20, 2025. The oldest animal at the San Diego Zoo was believed to be around 141 ...

The beloved Galápagos tortoise named Gramma that was the oldest resident of the San Diego Zoo at an estimated 141 years has ...

An occasional series where Mongabay founder Rhett Ayers Butler shares analysis, perspectives and story summaries. She moved ...

San Diego Zoo staff and animal lovers are reeling this week following the death of beloved Gramma the Galápagos tortoise, by ...

Families go wild when visiting San Diego, and when they throw in animal encounters, it gets even better. But here’s the catch: there are two big-name wildlife attractions in San Diego, and they’re ...

Gramma, a Galapagos tortoise that was the oldest living animal at the San Diego Zoo, has died at an estimated 141 years of ...